Chapter 55

Ecosystem ecology

-processes and interactions between species are important
- range from a micro to a large area
-ecologists

-Energy flow through ecosystem
-matter cycles within ecosystem

-abide by laws of physics

Ecologists

study the transformations of energy and matter within ecosystems

Ecosystem

community plus all the abiotic interactions associated with that community

Conservation of energy

- first and second law of thermodynamics
- energy enters ecosystem as solar radiation and is lost from organisms as heat
- energy conversions are not always efficient

first law of thermodynamics

Energy cannot be created or destroyed, only transformed

second law of thermodynamics

Every energy transfer or transformation increases the entropy of the universe.

Conservation of mass

- matter cannot be created or destroyed
-chemical elements are continually recycled within ecosystems
- in a forest, most nutrients enter as dust and carried out by water
-ecosystems are open, they absorb energy/mass and release heat/waste

Energy, mass and trophic levels

-autotrophs
-heterotrophs
-energy/nutrients pass from primary producers through many levels of consumers

Energy Budget

- primary production
- rate of primary production

Primary Production

fixation of energy by autotrophs in an ecosystem
and converts inorganic energy into organic energy
-aquatic ecosystems
-terrestrial ecosystems

Rate of Primary Production

amount of energy fixed over a given period of time
- gross and net

Gross

total amount of energy fixed by autotrophs

Net

Amount of energy leftover after needs are met

Aquatic Ecosystems

- light and nutrients control primary production
- depth of light penetration affects primary production in the photic zone of and ocean/lake
-nitrogen and phosphorus
-upwelling of nutrient rich waters
-large amounts of nutrients

Nitrogen and Phosphorous

nutrients that limit marine production

Upwelling of nutrient rich waters

high primary production

Large amounts of nutrients

- result in sewage runoff
- limits cyanobacterias growth
- reduce eutrophication

Terrestrial Ecosystems

- temp and moisture affect primary production
-primary production increase with moisture
-soil nutrients= limiting factor
-nitrogen and phosphorous= most common limiting nutrient

Energy

- transfer
-flow in temperate deciduous forest (Gosz)

Transfer

- 10% effecient between trophic levels
- apron. 0.1% of chemical energy fixed by photosynthesis reaches a tertiary consumer

Gosz's study of energy flow

- 99% of solar energy unavailable for use by second trophic level
-as energy losses between trophic levels, eventually there is insufficient energy to transport a population at a higher trophic level
- most food chains have 3-4 trophic levels with a maxim

Cycles

- biogeochemical
- water
- carbon
- phosphorus
-nitrogen

Biogeochemical

-Nutrient cycle
-gaseous carbon, oxygen, sulfur, and nitrogen occur in atmosphere and cycle globally
-less mobile elements

Nutrient Cycle

- that involves biotic and abiotic components
- transformation, movement and reuse of nutrients in ecosystems

Less mobile elements

phosphorous, potassium and calcium
- these cycle locally in terrestrial systems and broadly in aquatic

Water

- essential to all organisms
- liquid is primary physical phase water is used
- oceans contain 97% of biosphere water, 2% glaciers, 1% lakes/rivers
-moves by processes of evaporation, transpiration, condensation, precipitation and movement through groundw

Carbon

- important in all organic molecules and in atmosphere as carbon dioxide and methane
- moves in-between organisms and atmosphere as due to phototsythesis and respiration
-reservoirs

Reservoirs

fossil fuels, soils/sediments, solutes in ocean, plant/animal biomass, atmosphere, sedimentary rocks

CO2 is taken up and released through

photosynthesis and respiration
- volcanoes and burning fossil fuels contribute to CO2 in atmophere

Nitrogen

- can be released by fungi and bacteria during organism decomposition
- released as ammonium

Phosphorous

- important in ATP, RNA, DNA and cell membrane
- phosphate is most important inorganic form
- global phosphorous cycle does not include substantial atmospheric pool

Decomposition and nutrient cycling rates

next 6 questions

Decomposers play a key role in..

general pattern of chemical cycling

Rates that nutrients cycle vary greatly, mostly because of

decomposition

Rate of decomposition is controlled by

temperature, moisture, and nutrient availability

Rapid decomposition results in

relatively low levels of nutrients in the soil

Cold and wet ecosystems store large amounts of

undecomposed organic matter as decomposition rates are low

Cycling rates may be altered greatly by

human activities

Restoration Ecology

goal: return degraded ecosystems to a more natural state
- with time, bio communities can recover from many types of disturbances
- seeks to initiate of speed up the recovery of degraded ecosystems

Two key strategies in restoration

bioremediation and biological augmentation

Bioremediation

Use of living organisms such as prokaryotes, fungi, or plants to detoxify polluted ecosystems.
- can take up and metabolize toxic molecules

Augmentation

uses organisms to add essential materials to a degraded ecosystem
-long term objective of restoration

Long term objective of restoration

return ecosystem as much as possible to its predisturbance state