Ch. 9b Variation in Skeletal Muscle Contraction

motor units with smallest muscle fibers

activated first

motor units with largest muscle fibers

only activated when the most powerful contractions are necessary

what does muscle tension vary with

1. changes in stimulus voltage
2. changes in stimulus frequency

muscle twitch

single contraction followed by complete relaxation

latent period

muscle tension begins to increase, but no response seen on myogram

period of contraction

cross-bridges are active and muscle reaches peak tension

period of relaxation

reentry of calcium into SR as muscle tension decreases to zero

why do muscle twitch contractions vary

depends on fiber types contained in each muscle

what increases the number of motor units that are stimulated

strength of the stimulus (voltage)

recruitment

multiple motor unit summation

subthreshold stimulus

stimuli that produce no observable contractions

threshold stimulus

stimulus at which the first observable contraction occurs

maximal stimulus

strongest stimulus that produces increased contractile force

asynchonrous

usually occurs when all motor units are being stimulated
-helps to prevent or delay fatigue

increased stimulus frequency

can result in stronger muscle twitches

stimulus frequency between 10-20 stimuli per second

1. muscle twitches undergo complete relaxation
2. each subsequent twitch is stronger than the one before due to increased sarcoplasmic calcium levels
***referred to as TREPPE --> "Staircase effect

stimulus frequency between 20-40 stimuli per second

1. period of relaxation of each muscle twitch is interrupted by subsequent stimuli
2. causes successive contractions "piggy back" on one another, increasing tension
3. absolute refractory periods is always honored
***temporal wave summation which produces

stimulus frequency between 40-50 stimuli per second

1. period of relaxation of each muscle twitch is completely eliminated
2. produces one long, sustained contraction called Complete (fused) tetanus
3. rarely occurs in human body due to asynchronous nature of muscle fiber contraction

muscle twitches only produce what

internal tension

internal tension

when muscle fibers, contract, they pull on connective tissue sheaths (noncontractile components)

temporal summation produces what followed by what

produces internal tension followed by external tension

external tension

as sheaths become taut, they pull on the load (insertion); it takes longer to generate external tension than internal tension
**results in movement

length- tension relationship

amount of tension generated by muscle and force of its contraction depend on how stretched or contracted it was before stimulated

muscles between 60% and 175% of their resting length are able to generate the greatest muscle tension

true

a contracting muscle does not always shorten

true

muscle tension

force exerted by a contracting muscle on an object

load

weight of the object to be moved

isometric contractions

same measure"
1. muscle stays at a fixed length during contraction
2. muscle tension develops- doesn't overcome the weight of the load
3. cross-bridges generate force but filaments don't slide

passive force

generated by stretching the muscle and results from elastic recoil of the tissue itself
-due to Titin

active fore

generated during cross-bridge cycle

Total force

sum of passive and active forces

isotonic contractiosn

same tension
1. muscle tension sufficient to overcome the weight of the load
2. muscle will shorten and the load will be moved
3. filaments are sliding

2 types of isotonic contractions

1. concentric
2. eccentic

concentric

muscle shortens while doing work

eccentric

muscle is forced to elongate while contracting

what is the velocity and the duration of a contraction variable due to

1. more motor units
2. fiber type
3. weight of the load

more motor units results in what

faster, more prolonged contractions
-large muscles have more motor units and can generate a longer duration of contraction

fiber type

slow oxidative fibers can contract for longer periods, but fast glycolytic can often contract with a higher velocity

weight of the load

1. muscles contract fastest and most forcefully when there is no added load on them
2. increase load = increase latent period
3. increased load results in delayed contraction, and short contraction duration